Dr. John Hayden
Professor of Biology
Profile
Considered broadly, my lab investigates botanical biodiversity with two general aims, to improve our understanding of plant relationships (systematics) and to document plant diversity that exists within particular geographic locations (floristics). Our approach is largely structural, and work is done both in the field and in laboratory settings. The full range of plant morphology—from overall plant form, to macroscopic structures, to anatomical features discerned only with light or scanning electron microscope–is of interest. Recent investigations involve developmental floral anatomy to understand more thoroughly floral form, the cornerstone of plant systematics; such studies include Physalis and Jaltomata (Solanaceae), Flueggea (Phyllanthaceae) and Acalypha (Euphorbiaceae). Systematic work has often focused on plant family Euphorbiaceae (including its segregates Phyllanthaceae and Picrodendraceae) and inventory projects have been carried out in central Virginia and the Yucatan Peninsula of Mexico. Research is supported by greenhouse and herbarium collections.
Publications
Articles

Hayden, W. John. "Endless Symbioses Most Intricate." Sempervirens, Summer 2016, 12-13.

Hayden, W. John. "Seed Dispersal: A Tale of Two Species." Sempervirens, Spring 2016, 6-7.
Hayden, W. John. "There's Much Left to Learn: Clethra's Chromosomes. " Sempervirens, Fall 2015, 4-5.
Hayden, W. John. "When It Comes to Clethra: Roots Matter." Sempervirens, Summer 2015, 10-12.
Hayden, W. John. "2015 Virginia Wildflower of the Year: Sweet Pepperbush, Clethra alnifolia." Virginia Native Plant Society Brochure, 2015, 1-2.
Hayden, W. John. "Little Things Reveal the Big Picture." Sempervirens, Winter 2015, 4-5.
Hayden, W. John. "Native Orchids in Winter?." Sempervirens, Winter 2015, 5, 8.
Hayden, W. John. "Oh No! Something Is Eating My Coral Honeysuckle!" Bulletin of the Virginia Native Plant Society 33, no. 4 (August 2014): 1, 4.
Hayden, W. John. "Coral Honeysuckle Easy to Propagate with Cuttings." Bulletin of the Virginia Native Plant Society 33, no. 3 (Summer 2014): 3,5,8.
Hayden, W. John. "Humming Birds: Pollination Facts and Fancy" Bulletin of the Virginia Native Plant Society 33, no. 2 (Spring 2014): 1,5,8.
Hayden, W. John. "Two Honeysuckles: A Tale of Dr. Jekyll and Mr. Hyde." Bulletin of the Virginia Native Plant Society 33, no. 1 (February 2014): 1, 6,8.
W. John Hayden. 2014 Virginia Wildflower of the Year: Coral Honeysuckle, Lonicera sempervirens. Virginia Native Plant Society, 2014.
Hayden, W. J. 2013. Commelina benghalensis new to Virginia. Castanea 78: 329.
Hayden, W. John. "Redbud Seedpods Hold Surprises." Bulletin of the Virginia Native Plant Society 32, no. 5 (Autumn 2013): 1, 6
Hayden, W. John. "Redbuds Similar Around the World." Bulletin of the Virginia Native Plant Society 32, no. 3 (Summer 2013): 3,7.
Hayden, W. John. "Redbuds and Legumes Subfamilies." Bulletin of the Virginia Native Plant Society 32, no. 2 (Spring 2013): 1, 4-5.
Hayden, W. John. "Redbud Cauliflory: The Inside Story." Bulletin of the Virginia Native Plant Society 32, no. 1 (Winter 2013): 1, 3, 8.
Serdar, B., W. J. Hayden, & S. Terzioglu. 2008. Wood anatomy of Flueggea anatolica (Phyllanthaceae). IAWA Jour. 29: 303-310.
Terry*, M. A., & W. J. Hayden. 2007. Vascular Flora of Powhatan County, Virginia. Castanea 72: 138-158.
Terry, M., & Hayden, W.J. (2005) Vascular Plant Inventory of Powhatan County, Virginia. Virginia Journal of Science, 56.
Kostadinov, T., Hayden, S.M., & Hayden, W.J. (2004) Secondary Xylem of Dalechampia (Euphorbiaceae). Virginia Journal of Science, 55: 33-34.
Callaghan, J. A., & Hayden, W.J. (2004) The Helen Moyers Biocultural Reserve at Rancho Kiuic, Yucatan, Mexico: An Introduction to the Reserve and its Flora. Botany 2004 Scientific Meeting Abstracts, #663, p.169.
Kostadinov, T., Hayden, S.M., & Hayden, W.J. (2003) Correcting the Record: Wood of Dalechampia (Euphorbiaceae Subfamily Acalyphoideae) Lacks Laticifers. International Association of Wood Anatomists Journal, July 2003.
Hayden, W. J., & S. M. Hayden. 2000. Wood anatomy of Acalyphoideae (Euphorbiaceae). IAWA Journal 21: 213–235.
Hayden, W. J. 1999. Amanoa (Euphorbiaceae). Pp. 95-99, in: J. A. Steyermark et al., eds., Flora of the Venezuelan Guayana, Missouri Botanical Garden Press.
Terry*, M. & W. J. Hayden. 2005. Vascular Plants of Powhatan County, Virginia. Poster presented at the annual meeting of the Botanical Society of America, Austin, Texas.

Hayden, W. J. 2006. The Flora of Kaxil Kiuic, Yucatán, Mexico. Paper presented at the annual meeting of the Botanical Society of America, Chico, California.

Serdar, B., S.Terzioglu, & W. John Hayden. 2007. Wood anatomy of Flueggea anatolica (Phyllanthaceae). Paper presented at the Pan-American IAWA Meeting, San Luis Potosi, San Luis Potosi, Mexico, 20 July 2007.

Jahdi*, A., C. Marks, & W. John Hayden. 2007. Peculiar features of perforation plates of the Acalypha virginica complex (Euphorbiaceae). Paper presented at the Pan-American IAWA Meeting, San Luis Potosi, San Luis Potosi, Mexico, 20 July 2007.

Jahdi*, A., C. Marks, & W. J. Hayden. 2008. Intact perforation membranes in secondary xylem of the Acalypha virginica complex (Euphorbiaceae). Paper presented at the Botany 2008 annual meeting in Vancouver, British Columbia.

Hayden, W. J. & M. J. Drake*. 2010. Structure of foliar glands of Stizophyllum riparium (Bignoniaceae). Paper presented at the Botany 2010 annual meeting.

Dong*, D., P. M. Miller*, & W. J. Hayden. 2013. Floral Anatomy and Corolla Nectar Tubes in Physalis (Solanaceae). Poster presented, Botany 2013, New Orleans, LA.

*indicates UR student.
Additional Publications

Fagan*, C. & W. J. Hayden. 2016. Anatomy and pollination of cleistogamous flowers of Benghal Dayflower (Commelina benghalensis). Weed Science 64: 455—462. 

* denotes student coauthor

Hayden, W. John. "Upside-down Anthers of Clethra Stand Out." Sempervirens, Spring 2015, 1, 3.

Education
Ph.D., University of Maryland
Contact Information
(804) 289-8232
(804) 289-8233 (Fax)
Areas of Expertise
Anatomy, morphology, and systematics of vascular plants, especially family Euphorbiaceae, in the context of regional floristics, ethnobotany, systematic monographs, and revisions